The higher units of pine forests of Russia in connection with the general concept of vegetation classification of Northern Eurasia
https://doi.org/10.36305/2712-7788-2020-4-157-94-113
Abstract
A review of the classification of pine forests (dominated by two-needle pines of the subgenus Diploxylon) of Russia using the Braun-Blanquet approach was carried out. All diversity of pine forests was included in 9 higher units – vegetation classes, of which 3 classes represent typical communities dominated by pine species (Erico–Pinetea Horvat 1959, Koelerio glaucae–Pinetea sylvestris Ermakov class nova hoc loco, Junipero–Pinetea Rivas-Mart. 1965). In the remaining six classes (Vaccinio–Piceetea Br.-Bl. in Br.-Bl.et al. 1939, Carpino–Fagetea Jakucs et Passarge 1968, Quercetea pubescentis Doing-Kraft ex Scamoni et Passarge 1959, Brachypodio–Betuletea Ermakov et al. 1991, Rhytidio–Laricetea Korotkov et Ermakov 1999 and Quercetea mongolicae Song ex Krestov et al. 2006), pine forests are represented as special higher categories (alliances, orders) along with other types of zonal and non-zonal forest vegetation. The characteristics of the main higher units (classes, orders and alliances) are presented, as well as their diagnostic features and problems of classification of some categories are discussed. As a result of the syntaxonomic revision, the name of the class Pyrolo–Pinetea sylvestris Korneck 1974 was rejected, since the alliance Cytiso–Pinion sylvestris Krausch 1962 (the only alliance of order Pulsatillo–Pinetalia Oberd. in Oberd. et al. 1967) was proposed in the original article as provisional (3b). It is proposed to include xerophilous psammophilous pine forests of Europe and Western Siberia into a new class Koelerio glaucae–Pinetea sylvestris class nova hoc loco. The nomenclature type of the class (holotypus) is the order Koelerio glaucae–Pinetalia sylvestris Ermakov 1999. Diagnostic species of the class are Festuca beckeri, Gypsophylla altissima, G. paniculata, Jurinea cyanoides, Koeleria glauca, Oxytropis campanulata, Potentilla humifusa, Veronica spicata, Silene baschkirorum, S. chlorantha, Stipa pennata subsp. sabuletorum, Helichrysum arenarium, Centaurea arenaria, C. marschalliana, Achillea gebleri, Genista tinctoria, Cytisus ruthenica, C. borystenica, Hieracium pilosella, Dianthus arenarius.
About the Author
N. B. Ermakov
Никитский ботанический сад – Национальный научный центр РАН; Майкопский государственный технологический университет
Russian Federation
Russian Federation
References
1. Ермаков Н.Б. Синтаксономические и ботанико-географические особенности ксерофильных псаммофильных сосновых лесов Западно-Сибирской равнины // Флора и растительность Алтая. 1999. Т.4. № 1. С. 52-61. [Ermakov N.B. Syntaxonomic and plant-geographical peculiarities of xerophilous, psammophilous pine forests of West-Siberian plain. Flora and vegetation of the Altai. 1999. 4 (1):52-61.]
2. Ермаков Н.Б., Абдурахманова З. И., Потапенко И. Л. К проблеме синтаксономии сосновых лесов (Pinus sylvestris var. hamata) с участием бореальных флористических элементов в Дагестане (Северный Кавказ) // Turczaninowia . 2019. № 22 (4). С. 154-171. DOI: 10.14258/turczaninowia.22.4.16. [Ermakov N.B., Abdurakhmanova Z.I., Potapenko I.L. To the problem of syntaxonomy of pine forests (Pinus sylvestris var. hamata) with the participation of boreal floristic elements in Dagestan (North Caucasus). Turczaninowia. 2019. 22 (4):154-171]
3. Ермаков Н.Б., Королюк А.Ю., Лащинский Н.Н. Флористическая классификация мезофильных травяных лесов Южной Сибири. Новосибирск. 1991. 96 c. [Ermakov N.B., Korolyuk A.Yu., Lashchinsky N.N. Floristic classification of mesophilous grass forests of Southern Siberia. Novosibirsk. 1991. 96 p.]
4. Лапшина Е.Д. Растительность болот юго-востока Западной Сибири. Новосибирск. 2010. 186 c. [Lapshina E.D. Swamp vegetation of south-east of West Siberia. Novosibirsk. 2010. 186 p.]
5. Лащинский Н.Н., Королюк А.Ю., Лащинская Н.В. Классы Pyrolo-Pinetea и Vaccinio-Piceetea в растительном покрове Бурлинского ленточного бора // Растительный мир Азиатской России. 2017. № 26 (2). C. 63-84. [Lashchinsky N.N., Korolyuk A.Yu., Lashchinskaya N.V. Pyrolo-Pinetea and VaccinioPiceetea classes in the Burla ribbon pine forests vegetation cover. Plant life of Asian Russia. 2017. 26 (2):63-84]
6. Лащинский Н.Н., Королюк А.Ю., Тищенко М.П., Лащинская Н.В. Синтаксономия и пространственная структура растительности Бурлинского ленточного бора // Растительный мир Азиатской России. 2018. № 29 (1). C. 57-81. DOI:10.21782/RMAR1995-2449-2018-1(57–81) [Lashchinsky N.N., Korolyuk A.Yu., Tischenko M.P., Lashchinskaya N.V. Syntaxonomy and spatial structure of the Burla ribbon pine forests. Plant life of Asian Russia. 2018. 29 (1):57- 81]
7. Полякова М.А., Ермаков Н.Б. Классификация сосновых лесов боровых лент Минусинской межгорной котловины (Южная Сибирь) // Растительность России. 2008. № 13. C. 82-105. DOI: https://doi.org/10.31111/vegrus/2008.13.82 [Polyakova M.A., Ermakov N.B. Classification of pine forests on sandy deposits in the Minusinskaya intermountain basin (Southern Siberia). Vegetation of Russia. 2008. 13: 82- 105]
8. Широких П.С., Кунафин А.М., Бикбаев И.Г., Салихов Д.М., Мартыненко В.Б. Коррекция союза Dicrano-Pinion на основе синтаксономии и ординационного анализа // Известия Самарского центра Российской академии наук. 2013.№ 15 (3–1). С. 395-400. [Shirokikh P.S., Kunafin A.M., Bikbayev I.G., Salikhov D.M., Martynenko V.B. Correction of the alliance Dicrano-Pinion on the basis of syntaxonomy and ordination analysis. Izvestia of Samara Scientific Center of the Russian Academy of Sciences. 2013. 15 (3–1): 395-400]
9. Широких П.С., Мартыненко В.Б., Баишева Э.З., Бикбаев И.Г. Динамика растительности на вырубках Южно-Уральского региона: основные итоги исследований Уфимской геоботанической школы // Фиторазнообразие Восточной Европы. 2018. № XII (3). С. 17-30. DOI: 10.24411/2072-8816-2018-10023 [Shirokikh P.S., Martynenko V.B., Baisheva E.Z., Bikbayev I.G. Vegetation dynamic after clearings in the Southern Urals: the main results of studies of Ufa geobotanical school. Phytodiversity of Eastern Europe. 2018. XII (3):17-30]
10. Черепанов С.К. Сосудистые растения России и сопредельных государств. СанктПетербург. 1995. 990 c. [Cherepanov S.K. Vascular plants of Russia and neighboring countries. Saint-Petersburg. 1995. 990 p.]
11. Anenkhonov O.A., Chytry M. Syntaxonomy of vegetation of the Svyatoj Nos Peninsula, Lake Baikal 2. Forests and krummholz in comparison with other regions of northern Buryatia // Folia Geobotanica. 1998. Vol. 33 (1). P. 31-75. DOI: https://doi.org/10.1007/BF02914929
12. Barkman J.J., Moravec J., Rauschert S. Code of phytosociological nomenclature // Vegetatio. 1976. Vol. 32. P. 131-185. https://doi.org/10.1007/BF02095917
13. Berg C., Dengler J., Abdank A., Isermann M. (eds.) Die Pflanzengesellschaften Mecklenburg-Vorpommerns und ihre Gefahrdung. Textband. 2004. Weissdorn. Jena. 606 p.
14. Bergmeier E., Walentowski H., Güngöroğlu C. Turkish Forest Habitat Types – An Annotated Conspectus Based on the EU Habitats Directive with Suggestions for an Upgrade. In: Practicability of EU Natura 2000 Concept in the Forested Areas of Turkey. Ankara: Forester’s Association Publisher. 2019. P. 134-292.
15. Coban S., Willner W. Numerical classification of the forest vegetation in the Western Euxine Region of Turkey // Phytocoenologia. 2019. Vol. 49 (1). P. 71-106. DOI: 10.1127/phyto/2018/0274
16. Ermakov, N., Cherosov, M., Gogoleva, P. Classification of ultracontinental boreal forests in Central Yakutia // Folia Geobotanica. 2002. Vol. 37 (4). P. 419-440. https://doi.org/10.1007/BF02803256
17. Ermakov N., Morozova O. Syntaxonomical survey of boreal oligotrophic pine forests in northern Europe and Western Siberia // Applied Vegetation Science. 2011. Vol. 14 (4). P. 524-536. DOI: 10.1111/j.1654-109X.2011.01155.x
18. Ignatov M. S., Afonina O. M., Ignatova E. A., Abolina A., Akatova T. V., Baisheva E. Z., Bardunov L. V., Baryakina E. A., Belkina O. A., Bezgodov A. G., Boychuk M. A., Cherdantseva V. Y., Czernyadjeva I. V., Doroshina G. Y., Dyachenko A. P., Fedosov V. E., Goldberg I. L., Ivanova E. I., Jukoniene I., Kannukene L., Kazanovsky S. G., Kharzinov Z. K., Kurbatova L. E., Maksimov A. I., Mamatkulov U. K., Manakyan V. A., Maslovsky O. M., Napreenko M. G., Otnyukova T. N., Partyka L. Y., Pisarenko O. Y., Popova N. N., Rykovsky G. F., Tubanova D. Y., Zheleznova G. V., Zolotov V. I. Check-list of mosses of East Europe and North Asia // Arctoa. 2006. Vol. 15. P. 1-130. DOI:10.15298/arctoa.15.01
19. Korneck D. Xerothermvegetation in Rheinland-Pflaz und Nachbargebieten // Schriftenreihe fur Vegetationskunde. 1974. Vol. 7. P. 1-196.
20. Korotkov K., Morozova O., Belanovskaja E. The USSR Vegetation Syntaxa Prodromus. Moscow: Published by Dr. G. E. Vilchek, 1991. 346 p.
21. Korzhenevsky V.V. Pinus pallasiana forests in the Crimea // Укр. фiтосоцiол. зб. Сер. А. Kиïв. 1998. № 1 (9). С. 78- 97.
22. Krausch H-D. Der Sandnelken-Kiefernwald an seiner Westgrenze in Brandenburg // Mitteilungen der Floristisch-soziologischen Arbeitsgemeinschaft (alte Serie). 1962. Vol. 9. P. 141-144.
23. Krestov P.V., Song J.-S., Nakamura Yu., Verkholat V.P. A phytosociological survey of the deciduous temperate forests of mainland Northeast Asia // Phytocoenologia. 2006. Vol. 36 (1). P. 77-150. DOI: 10.1127/0340-269X/2006/0036-0077
24. Libbert W. Die Vegetationseinheiten der Neumarkischen Staubeckenlandschaft // Verhandlungen des Botanischen Vereins der Provinz Brandenburg. 1933. Vol. 74. P. 229-348.
25. Matuszkiewicz W., Matuszkiewicz J. Przeglad fitosocjologiczny zbiorowisk lesnych Polski. Cz. 2. Bory sosnowe // Phytocoenosis. Biuletyn Fitosocjologiczny. 1973. Vol. 2. P. 273-356.
26. Mucina L., Grabherr G., Wallnofer S. (eds). The plant communities of Austria. № 3. Forest and shrub communities. Jena: Gustav Fischer, 1993. 376 p.
27. Mucina L., Bultmann H., Dierssen K., Theurillat J.P., Raus T., Carni A., Sumberova K., Willner W., Dengler J., Garcia R., Chytry M., Hajek M., Di Pietro R., Yakushenko D., Pallas J., Daniels F., Bergmeier E., Guerra A., Ermakov N., Valachovic M., Schaminee J., Lysenko T., Didukh Y., Pignatti S., Rodwell J., Capelo J., Weber H., Solomeshch A., Dimopoulos P., Aguiar C., Hennekens S., Tichy L. Vegetation of Europe: hierarchical floristic classification systemof vascular plant, bryophyte, lichen, and algal communities // Applied Vegetation Science. 2016. Vol. 19 (1). P. 3-264. Doi: 10.1111/avsc.12257.
28. Oberdorfer E. Süddeutsche Pflanzengesellschaften. Jena: Gustav Fischer Verlag, 1957. 564 p.
29. Weber H. E., Moravec J., Theurillat J.-P. International code of phytosociological nomenclature. 3rd edition // Journal of Vegetation Science. 2000. Vol.11 (5). P. 739-768. DOI: 10.2307/3236580
30. Westhoff V., van der Maarel E. The Braun-Blanquet approach. In: Whittaker R.H. (ed.) Handbook of vegetation science. Part 5. Classification and ordination of communities. Junk, The Hague. 1973. P. 617-726.
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Ermakov N.B. The higher units of pine forests of Russia in connection with the general concept of vegetation classification of Northern Eurasia. Plant Biology and Horticulture: theory, innovation. 2020;1(157):94-113. (In Russ.) https://doi.org/10.36305/2712-7788-2020-4-157-94-113
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