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<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="ru"><front><journal-meta><journal-id journal-id-type="publisher-id">scbook</journal-id><journal-title-group><journal-title xml:lang="ru">Биология растений и садоводство: теория, инновации</journal-title><trans-title-group xml:lang="en"><trans-title>Plant Biology and Horticulture: theory, innovation</trans-title></trans-title-group></journal-title-group><issn pub-type="ppub">2712-7788</issn><publisher><publisher-name>Federal State Funded Institution of Science “The Labour Red Banner Order Nikitsky Botanical Gardens – National scientific Center of the RAS”</publisher-name></publisher></journal-meta><article-meta><article-id custom-type="elpub" pub-id-type="custom">scbook-851</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>БОТАНИКА И ЭКОЛОГИЯ РАСТЕНИЙ</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>BOTANY AND PLANT ECOLOGY</subject></subj-group></article-categories><title-group><article-title>Подходы к определению редкости папоротникообразных (включая плаунообразные) Социалистической Республики Вьетнам</article-title><trans-title-group xml:lang="en"><trans-title>Approaches to determining the rarity of ferns (including Lycopodiophyta) for the Socialist Republic of Vietnam</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Калюжный</surname><given-names>С. С.</given-names></name><name name-style="western" xml:lang="en"><surname>Kalyuzhny</surname><given-names>S. S.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Сергей Сергеевич Калюжный</p><p>97022; ул. Профессора Попова, д. 2, литера В; Санкт-Петербург</p></bio><bio xml:lang="en"><p>97022; 2 Professora Popova Street, Building B; Saint-Petersburg</p></bio><email xlink:type="simple">fernbaikal@yandex.ru</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Шеметова</surname><given-names>И. С.</given-names></name><name name-style="western" xml:lang="en"><surname>Shemetova</surname><given-names>I. S.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Инна Сергеевна Шеметова</p><p>664038; пос. Молодежный, здание 1/1; Иркутская область</p></bio><bio xml:lang="en"><p>664038; Molodezhny settlement, building 1/1; Irkutsk region</p></bio><email xlink:type="simple">inna198410@mail.ru</email><xref ref-type="aff" rid="aff-2"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Каленчук</surname><given-names>Т. В.</given-names></name><name name-style="western" xml:lang="en"><surname>Kalenchuk</surname><given-names>T. V.</given-names></name></name-alternatives><bio xml:lang="ru"><p>225710; ул. Днестровской флотилии д. 23; Брестская область; Пинск</p></bio><bio xml:lang="en"><p>225710; 23 Dnestrovskoy Flotilii Street; Brest Region; Pinsk</p></bio><email xlink:type="simple">chrysanthemum@list.ru</email><xref ref-type="aff" rid="aff-3"/></contrib></contrib-group><aff-alternatives id="aff-1"><aff xml:lang="ru"><institution>ФГБНУ Ботанический институт им. В.Л. Комарова –&#13;
Ботанический сад Петра Великого</institution><country>Россия</country></aff><aff xml:lang="en"><institution>FGBNU V.L. Komarov Botanical Institute – Peter the Great Botanical Garden</institution><country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-2"><aff xml:lang="ru"><institution>ФГБОУ ВО Иркутский государственный аграрный университет</institution><country>Россия</country></aff><aff xml:lang="en"><institution>Irkutsk State Agrarian University</institution><country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-3"><aff xml:lang="ru"><institution>УО ПолесГУ</institution><country>Беларусь</country></aff><aff xml:lang="en"><institution>UO PolesSU</institution><country>Belarus</country></aff></aff-alternatives><pub-date pub-type="collection"><year>2025</year></pub-date><pub-date pub-type="epub"><day>22</day><month>03</month><year>2026</year></pub-date><volume>0</volume><issue>4 (177)</issue><fpage>43</fpage><lpage>55</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Калюжный С.С., Шеметова И.С., Каленчук Т.В., 2026</copyright-statement><copyright-year>2026</copyright-year><copyright-holder xml:lang="ru">Калюжный С.С., Шеметова И.С., Каленчук Т.В.</copyright-holder><copyright-holder xml:lang="en">Kalyuzhny S.S., Shemetova I.S., Kalenchuk T.V.</copyright-holder><license xml:lang="ru" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>Данная работа распространяется под лицензией Creative Commons Attribution 4.0.</license-p></license><license xml:lang="en" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://scbook.elpub.ru/jour/article/view/851">https://scbook.elpub.ru/jour/article/view/851</self-uri><abstract><p>   Определение редких и находящихся под угрозой исчезновения видов растений является основополагающим требованием для программы любого уровня по сохранению биоразнообразия. По данным IUCN в мире 12,5 % высших сосудистых растений находятся под угрозой исчезновения. Социалистическая Республика Вьетнам (СРВ) является страной с высоким уровнем биологического разнообразия, где произрастает около 12000-15000 видов сосудистых растений, включая 1132 вида папоротникообразных растений. В пределах этого региона были разработаны различные нормативные документы относительно статуса сохранения видов растений, находящихся под угрозой исчезновения, а также были изданы Красные книги. Было выпущено два указа Министерством сельского хозяйства и развития сельских районов (MARD) и Министерством природных ресурсов и окружающей среды (MONRE). Последний из них – Указ 84/2021 (2021), в котором перечислены две группы видов, IA и IIA, из растений и животных, произрастающих и отмеченных на территории СРВ, запрещенных для сбора/ловли и коммерческого использования. Папоротникообразные растения можно найти только в списке IIA: Huperzia serrata (Thunb.) Trevis., все виды Cyathea ssp. (после таксономических изменений это роды Alsophila R. Br – 5 видов, Gymnosphaera Blume – 3, Sphaeropteris Bernh. – 3), Cibotium barometz (L.) J.Sm., Drynaria bonii Christ, D. roosii Nakaike. Рассматривая категории редкости, а также встречаемости/распространенности папоротникообразных растений на территории СРВ необходимо отметить, что часть видов отмечена только по гербарным образцам. Из-за неизученности птеридофлоры, многие виды, приводятся в данной статье впервые для региона такие как Adiantum membranifolium S.Linds. &amp; Suksathan, Anisocampium cuspidatum (Mett. ex Bedd.) Yea C. Liu, W. L. Chiou &amp; M. Kato. Работая в полевых условиях на севере СРВ, были обнаружены эндемики Филиппин: редчайший Diplazium pseudocyatheifolium Rosenst. и Microsorum sarawakense (Baker) Holttum. Некоторые виды, были описаны только с гербарных образцов: Leptochilus heterophyllus (S. K. Wu &amp; P. K. Lôc) Kalyuzny (syn. Kontumia heterophylla S.K.Wu &amp; P.K.Lôc), который приводился как узколокальный эндемик для провинции Контум. Многие виды, которые приводились для Китайской Народной Республики (КНР), также были обнаружены: Athyrium erythropodum Hayata, Cyrtomium serratum Ching &amp; K. H. Shing, Coniogramme emeiensis Ching &amp; K. H.Shing и др. Имеется много документов, регламентирующих различную деятельность, законы об ООПТ, в том числе, Красная книга. Многие законы не работают на местном уровне, Красная книга выходит нерегулярно (первое издание опубликовано в 1996 году, переиздание в 2007). В настоящее время, с учетом новых данных, необходимы пересмотр и существенные изменения последнего издания. Следует отметить отсутствие взаимодействия со стороны СРВ с IUCN (Международный союз охраны природы и природных ресурсов), не ведутся и не составляются списки редких видов для международной охраны Красный список МСОП (IUCN – (Red List, 2012). Но лесное законодательство развивается и появляются новые законы, ограничивающие хозяйственную деятельность.</p></abstract><trans-abstract xml:lang="en"><p>   Identifying rare and endangered plant species is a fundamental requirement for any level of biodiversity conservation program. According to the IUCN, 12.5 % of higher vascular plants worldwide are threatened with extinction. The Socialist Republic of Vietnam (SRV) is a country with enormous biodiversity, where noted approximately 12000-15000 vascular plant species, including 1132 fern species. SRV has developed various regulations regarding the conservation status of threatened plant species and published Red Data Book. Two decrees have been issued by the Ministry of Agriculture and Rural Development (MARD) and the Ministry of Natural Resources and Environment (MONRE). The most recent of these is Decree 84/2021 (2021), which lists two species groups, IA and IIA, of plants and animals growing and recorded in SRV, prohibited for collection/catching and commercial use. Ferns can be found only in List IIA: Huperzia serrata (Thunb.) Trevis., all species of Cyathea ssp. (after taxonomic changes these are the genera Alsophila R. Br, Gymnosphaera Blume, Sphaeropteris Bernh.), Cibotium barometz (L.) J. Sm., Drynaria bonii Christ, D. roosii Nakaike. Considering the categories of rarity, as well as the occurrence/distribution of ferns in the territory of the SRV, it should be noted that some species are noted only from herbarium specimens. Due to the practical lack of study of the pteridoflora, many species are presented by us for the first time within the region, such as Adiantum membranifolium S. Linds. &amp; Suksathan, Anisocampium cuspidatum (Mett. ex Bedd.) Yea C. Liu, W.L. Chiou &amp; M. Kato. While working in the field in northern Vietnam, we found endemic species from Philippine: the extremely rare Diplazium pseudocyatheifolium Rosenst., and, as it turned out, Microsorum sarawakense (Baker) Holttum, which is well represented outside the locus classicus. Some species were described only from herbarium specimens: Leptochilus heterophyllus (S. K. Wu &amp; P. K. Lôc) Kalyuzny (syn. Kontumia heterophylla S. K. Wu &amp; P. K. Lôc), which was cited as a local endemic for Kontum Province. We also found many species previously reported for the People's Republic of China (PRC): Athyrium erythropodum Hayata, Cyrtomium serratum Ching &amp; K. H. Shing, Coniogramme emeiensis Ching &amp; K. H. Shing, and others. As can be seen from the above review, there are numerous documents regulating various activities, including laws on protected areas and the Red Book. However, many laws are ignored by local populations; the Red Book is published very irregularly (the first was published in 1996, the second in 2007), and there is currently no more recent edition. Unfortunately, it should be noted that there is absolutely no interaction between the PRC and the IUCN, and lists of rare species for international protection (Red List) are not maintained or compiled. However, forestry legislation is evolving, and new laws are emerging that restrict economic activities.</p></trans-abstract><kwd-group xml:lang="ru"><kwd>папоротникообразные растения</kwd><kwd>Diplazium pseudocyatheifolium Rosenst.</kwd><kwd>Adiantum membranifolium S. Linds. &amp; Suksathan</kwd><kwd>редкие и находящиеся под угрозой исчезновения папоротники</kwd><kwd>идентификация</kwd><kwd>Социалистическая Республика Вьетнам</kwd></kwd-group><kwd-group xml:lang="en"><kwd>ferns</kwd><kwd>Diplazium pseudocyatheifolium Rosenst.</kwd><kwd>Adiantum membranifolium S. Linds. &amp; Suksathan</kwd><kwd>rare and endangered ferns</kwd><kwd>identification</kwd><kwd>Socialist Republic of Vietnam</kwd></kwd-group><funding-group><funding-statement xml:lang="ru">Исследование выполнено в рамках государственного задания «История создания, состояние, потенциал развития живых коллекций растений Ботанического сада Петра Великого БИН РАН» 124020100075-2</funding-statement><funding-statement xml:lang="en">The research was carried out within the framework of the state assignment "The history of the creation, condition, and development potential of the living plant collections of the Peter the Great Botanical Garden." Great BIN RAN" 124020100075-2</funding-statement></funding-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Мочалов А.С. Птеридофлора как объект изучения // Вестник Курганского государственного университета. Сер. Естественные науки. 2013. Вып. 6. № 3 (30). С. 10-14.</mixed-citation><mixed-citation xml:lang="en">Mochalov A.S. Pteridoflora kak ob"ekt izucheniya // Vestnik Kurganskogo gosudarstvennogo universiteta. Ser. Estestvennye nauki. 2013. 6(3) (30):10-14.</mixed-citation></citation-alternatives></ref><ref id="cit2"><label>2</label><citation-alternatives><mixed-citation xml:lang="ru">Тахтаджян А.Л. Растения в системе организмов. Жизнь растений Т. 1. М.: Наука, 1974. С. 49-57.</mixed-citation><mixed-citation xml:lang="en">Tahtadzhyan A.L. Rasteniya v sisteme organizmov. Zhizn' rastenij T. 1. M.: Nauka, 1974. P. 49-57.</mixed-citation></citation-alternatives></ref><ref id="cit3"><label>3</label><citation-alternatives><mixed-citation xml:lang="ru">Толмачев А.И. Введение в географию растений. Л.: Изд-во ЛГУ, 1974. 244 с.</mixed-citation><mixed-citation xml:lang="en">Tolmachev A.I. Vvedenie v geografiyu rastenij. L.: Izd-vo LGU, 1974. 244 p.</mixed-citation></citation-alternatives></ref><ref id="cit4"><label>4</label><citation-alternatives><mixed-citation xml:lang="ru">A community-derived classification for extant lycophytes and ferns // Journal of systematics and evolution. PPG I. The Pteridophytes Phylogeny Group. 2016. Vol. 54 (6). P. 563-603. DOI: 10.1111/jse.12229</mixed-citation><mixed-citation xml:lang="en">A community-derived classification for extant lycophytes and ferns // Journal of systematics and evolution. PPG I. The Pteridophytes Phylogeny Group. 2016. Vol. 54 (6). P. 563-603. DOI: 10.1111/jse.12229</mixed-citation></citation-alternatives></ref><ref id="cit5"><label>5</label><citation-alternatives><mixed-citation xml:lang="ru">Adam Zajqc, Maria Zajqc. Distribution Atlas of Vascular Plants in Poland (ATPOL). Polish Academy of Sciences, 1992. 32 p.</mixed-citation><mixed-citation xml:lang="en">Adam Zajqc, Maria Zajqc. Distribution Atlas of Vascular Plants in Poland (ATPOL). Polish Academy of Sciences, 1992. 32 p.</mixed-citation></citation-alternatives></ref><ref id="cit6"><label>6</label><citation-alternatives><mixed-citation xml:lang="ru">Averyanov L.V., Nguyen K.S., Tran T.H., Averyanova A.L., Maisak T.V. &amp; Nguyen H.T. Plant diversity, flora and vegetation of Bat Dai Son Mountain area, northern Vietnam. Saint-Petersburg: Strata, 2020. 130 p.</mixed-citation><mixed-citation xml:lang="en">Averyanov L.V., Nguyen K.S., Tran T.H., Averyanova A.L., Maisak T.V. &amp; Nguyen H.T. Plant diversity, flora and vegetation of Bat Dai Son Mountain area, northern Vietnam. Saint-Petersburg: Strata, 2020. 130 p.</mixed-citation></citation-alternatives></ref><ref id="cit7"><label>7</label><citation-alternatives><mixed-citation xml:lang="ru">Bakgh – Comeau Y.S. Risk indexing rating of threatened ferns in Trinidad and Tobago. In: J.M. Camus, M. Gibby &amp; R.J. John (Edu) // Pteridology in perspective: Proc. Holttum memorial Pterid. Sympo. Kew. 1999. P. 139-152.</mixed-citation><mixed-citation xml:lang="en">Bakgh – Comeau Y.S. Risk indexing rating of threatened ferns in Trinidad and Tobago. In: J.M. Camus, M. Gibby &amp; R.J. John (Edu) // Pteridology in perspective: Proc. Holttum memorial Pterid. Sympo. Kew. 1999. P. 139-152.</mixed-citation></citation-alternatives></ref><ref id="cit8"><label>8</label><citation-alternatives><mixed-citation xml:lang="ru">Benniamin A., Irudayaraj V., Manickam V.S. How to Identify Rare and Endangered Ferns and Fern Allies // Ethnobotanical Leaflets. 2008. Vol. 12. P. 108-117.</mixed-citation><mixed-citation xml:lang="en">Benniamin A., Irudayaraj V., Manickam V.S. How to Identify Rare and Endangered Ferns and Fern Allies // Ethnobotanical Leaflets. 2008. Vol. 12. P. 108-117.</mixed-citation></citation-alternatives></ref><ref id="cit9"><label>9</label><citation-alternatives><mixed-citation xml:lang="ru">Dang N.V., Mai T.D., Chu H.C., Huy T.D., Kinh N.H. Forestry in Vietnam (1945-2000), development progress and experienced lessons. Agricultural Publishing House, 2001.</mixed-citation><mixed-citation xml:lang="en">Dang N.V., Mai T.D., Chu H.C., Huy T.D., Kinh N.H. Forestry in Vietnam (1945-2000), development progress and experienced lessons. Agricultural Publishing House, 2001.</mixed-citation></citation-alternatives></ref><ref id="cit10"><label>10</label><citation-alternatives><mixed-citation xml:lang="ru">Darlington C.D. Recent Advances in Cytology. London: J. &amp; A. Churchill, 1965. 768 p.</mixed-citation><mixed-citation xml:lang="en">Darlington C.D. Recent Advances in Cytology. London: J. &amp; A. Churchill, 1965. 768 p.</mixed-citation></citation-alternatives></ref><ref id="cit11"><label>11</label><citation-alternatives><mixed-citation xml:lang="ru">Decree 156/2018/ND-CP. On enforcement of a number of articles of the law on forestry. Hanoi. https://vanban.chinhphu.vn/?pageid=27160&amp;docid=195435. 2018.</mixed-citation><mixed-citation xml:lang="en">Decree 156/2018/ND-CP. On enforcement of a number of articles of the law on forestry. Hanoi. https://vanban.chinhphu.vn/?pageid=27160&amp;docid=195435. 2018.</mixed-citation></citation-alternatives></ref><ref id="cit12"><label>12</label><citation-alternatives><mixed-citation xml:lang="ru">Decree 84/2021/ND-CP. 2021. On management of endangered, precious and rare forest plants and animals and implementation of the Convention on International Trade in Endangered Species of Wild Fauna and Flora. Hanoi. https://vanban.chinhphu.vn/default.aspx?pageid=27160&amp;docid=204157.</mixed-citation><mixed-citation xml:lang="en">Decree 84/2021/ND-CP. 2021. On management of endangered, precious and rare forest plants and animals and implementation of the Convention on International Trade in Endangered Species of Wild Fauna and Flora. Hanoi. https://vanban.chinhphu.vn/default.aspx?pageid=27160&amp;docid=204157.</mixed-citation></citation-alternatives></ref><ref id="cit13"><label>13</label><citation-alternatives><mixed-citation xml:lang="ru">Dinerstein E. Joshi A.R., Vynne C. et al. A “Global Safety Net” to reverse biodiversity loss and stabilize Earth’s climate // Science Advances. 2020. Vol. 6 (36). eabb2824.</mixed-citation><mixed-citation xml:lang="en">Dinerstein E. Joshi A.R., Vynne C. et al. A “Global Safety Net” to reverse biodiversity loss and stabilize Earth’s climate // Science Advances. 2020. Vol. 6 (36). eabb2824.</mixed-citation></citation-alternatives></ref><ref id="cit14"><label>14</label><citation-alternatives><mixed-citation xml:lang="ru">Do Q., Bravo F., Sierra-de-Grado R., et al. Global biodiversity-related conventions on facilitating biodiversity conservation in Vietnam // Forest and Society. 2022. Vol. 6 (2). P. 489-502. DOI: 10.24259/fs.v6i2.14473</mixed-citation><mixed-citation xml:lang="en">Do Q., Bravo F., Sierra-de-Grado R., et al. Global biodiversity-related conventions on facilitating biodiversity conservation in Vietnam // Forest and Society. 2022. Vol. 6 (2). P. 489-502. DOI: 10.24259/fs.v6i2.14473</mixed-citation></citation-alternatives></ref><ref id="cit15"><label>15</label><citation-alternatives><mixed-citation xml:lang="ru">Dolores Ferrer-Castan, Ole R. Vetaas. Pteridophyte richness, climate and topography in the Iberian Peninsula // Global Ecology and Biogeography. 2004. Vol. 14(2). P. 155–165. DOI: 10.1111/J.1466-822X.2004.00140.X.</mixed-citation><mixed-citation xml:lang="en">Dolores Ferrer-Castan, Ole R. Vetaas. Pteridophyte richness, climate and topography in the Iberian Peninsula // Global Ecology and Biogeography. 2004. Vol. 14(2). P. 155–165. DOI: 10.1111/J.1466-822X.2004.00140.X.</mixed-citation></citation-alternatives></ref><ref id="cit16"><label>16</label><citation-alternatives><mixed-citation xml:lang="ru">Dong‐Mei Jin, Xi‐Le Zhou, Harald Schneider, Hong‐Jin Wei, Hong‐Yu Wei, Yue‐Hong Yan. Functional traits: Adaption of ferns in forest // Journal of Systematics and Evolution. 2021. Vol. 59(5). P. 1040-1050. DOI: 10.1111/jse.12669</mixed-citation><mixed-citation xml:lang="en">Dong‐Mei Jin, Xi‐Le Zhou, Harald Schneider, Hong‐Jin Wei, Hong‐Yu Wei, Yue‐Hong Yan. Functional traits: Adaption of ferns in forest // Journal of Systematics and Evolution. 2021. Vol. 59(5). P. 1040-1050. DOI: 10.1111/jse.12669</mixed-citation></citation-alternatives></ref><ref id="cit17"><label>17</label><citation-alternatives><mixed-citation xml:lang="ru">Fawcett S. &amp; Smith A.R. A generic classification of the Thelypteridaceae. Sida, Bot. Misc. 59. BRIT Press, Fort Worth Botanic Garden. Botanical Research Institute of Texas, U.S.A., 2021. 102 p.</mixed-citation><mixed-citation xml:lang="en">Fawcett S. &amp; Smith A.R. A generic classification of the Thelypteridaceae. Sida, Bot. Misc. 59. BRIT Press, Fort Worth Botanic Garden. Botanical Research Institute of Texas, U.S.A., 2021. 102 p.</mixed-citation></citation-alternatives></ref><ref id="cit18"><label>18</label><citation-alternatives><mixed-citation xml:lang="ru">Forest Department. Forest development in period of 2006-2010 // Journal of Agriculture and Rural Development. 2004. Vol. 12. P. 13-17.</mixed-citation><mixed-citation xml:lang="en">Forest Department. Forest development in period of 2006-2010 // Journal of Agriculture and Rural Development. 2004. Vol. 12. P. 13-17.</mixed-citation></citation-alternatives></ref><ref id="cit19"><label>19</label><citation-alternatives><mixed-citation xml:lang="ru">Hong Qian, Michael Kessler, Ole R. Vetaas. Pteridophyte species richness in the central Himalaya is limited by cold climate extremes at high elevations and rainfall seasonality at low elevations // Ecology and Evolution. 2022. Vol. 12(5). Article e8958. DOI: 10.1002/ece3.8958.</mixed-citation><mixed-citation xml:lang="en">Hong Qian, Michael Kessler, Ole R. Vetaas. Pteridophyte species richness in the central Himalaya is limited by cold climate extremes at high elevations and rainfall seasonality at low elevations // Ecology and Evolution. 2022. Vol. 12(5). Article e8958. DOI: 10.1002/ece3.8958.</mixed-citation></citation-alternatives></ref><ref id="cit20"><label>20</label><citation-alternatives><mixed-citation xml:lang="ru">Hung T.V. Forestry research for the coming time // Journal of Agriculture and Rural Development. 2004. Vol. 12. P. 26-29.</mixed-citation><mixed-citation xml:lang="en">Hung T.V. Forestry research for the coming time // Journal of Agriculture and Rural Development. 2004. Vol. 12. P. 26-29.</mixed-citation></citation-alternatives></ref><ref id="cit21"><label>21</label><citation-alternatives><mixed-citation xml:lang="ru">Brondizio E.S., Settele J., Diaz S., Ngo H.T. IPBES, Global assessment report of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services. (Eds). IPBES secretariat, Bonn, Germany, 2019. 1144 p.</mixed-citation><mixed-citation xml:lang="en">Brondizio E.S., Settele J., Diaz S., Ngo H.T. IPBES, Global assessment report of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services. (Eds). IPBES secretariat, Bonn, Germany, 2019. 1144 p.</mixed-citation></citation-alternatives></ref><ref id="cit22"><label>22</label><citation-alternatives><mixed-citation xml:lang="ru">Irudayaraj V., Manickam V.S., Johnson M. Vivipary, a rare and evolutionary important phenomenon in a rare homosporous fern Grammitis medialis from the Western Ghats // Current Science. 2003. Vol. 85(12). P. 1666-1667.</mixed-citation><mixed-citation xml:lang="en">Irudayaraj V., Manickam V.S., Johnson M. Vivipary, a rare and evolutionary important phenomenon in a rare homosporous fern Grammitis medialis from the Western Ghats // Current Science. 2003. Vol. 85(12). P. 1666-1667.</mixed-citation></citation-alternatives></ref><ref id="cit23"><label>23</label><citation-alternatives><mixed-citation xml:lang="ru">IUCN Red List categories and criteria, version 3.1, second edition. Gland and Cambridge, 2012. 32 p.</mixed-citation><mixed-citation xml:lang="en">IUCN Red List categories and criteria, version 3.1, second edition. Gland and Cambridge, 2012. 32 p.</mixed-citation></citation-alternatives></ref><ref id="cit24"><label>24</label><citation-alternatives><mixed-citation xml:lang="ru">Kalyuzhny S.S., Plugatar Y.V., Shalimov A.P. Modern taxonomic list of pteridophytes (including Lycopodiopsida class) for the Socialist Republic of Vietnam // Indian Fern Journal 2024. Vol. 41 (1). P. 53–75.</mixed-citation><mixed-citation xml:lang="en">Kalyuzhny S.S., Plugatar Y.V., Shalimov A.P. Modern taxonomic list of pteridophytes (including Lycopodiopsida class) for the Socialist Republic of Vietnam // Indian Fern Journal 2024. Vol. 41 (1). P. 53–75.</mixed-citation></citation-alternatives></ref><ref id="cit25"><label>25</label><citation-alternatives><mixed-citation xml:lang="ru">Law on Forestry. 2017. The National Assembly of Vietnam. Hanoi. https://vanban.chinhphu.vn/?pageid=27160&amp;docid=192329.</mixed-citation><mixed-citation xml:lang="en">Law on Forestry. 2017. The National Assembly of Vietnam. Hanoi. https://vanban.chinhphu.vn/?pageid=27160&amp;docid=192329.</mixed-citation></citation-alternatives></ref><ref id="cit26"><label>26</label><citation-alternatives><mixed-citation xml:lang="ru">Lindsay S., Middleton D.J. Ferns of Thailand, Laos and Cambodia (2016). Available at: http://rbgweb2.rbge.org.uk/thaiferns (accesd 23. 09. 2025).</mixed-citation><mixed-citation xml:lang="en">Lindsay S., Middleton D.J. Ferns of Thailand, Laos and Cambodia (2016). Available at: http://rbgweb2.rbge.org.uk/thaiferns (accesd 23. 09. 2025).</mixed-citation></citation-alternatives></ref><ref id="cit27"><label>27</label><citation-alternatives><mixed-citation xml:lang="ru">Lung N.N. The status of forest resources in Vietnam: matter of environment, economy, society and resolutions // Journal of Agriculture and Rural Development. 2001. Vol. 12. P. 891-893.</mixed-citation><mixed-citation xml:lang="en">Lung N.N. The status of forest resources in Vietnam: matter of environment, economy, society and resolutions // Journal of Agriculture and Rural Development. 2001. Vol. 12. P. 891-893.</mixed-citation></citation-alternatives></ref><ref id="cit28"><label>28</label><citation-alternatives><mixed-citation xml:lang="ru">Lycopodiopsida in GBIF Secretariat. GBIF Backbone Taxonomy. Checklist dataset doi: 10.15468/39omei accessed via GBIF.org on 2024-11-02. 2023.</mixed-citation><mixed-citation xml:lang="en">Lycopodiopsida in GBIF Secretariat. GBIF Backbone Taxonomy. Checklist dataset doi: 10.15468/39omei accessed via GBIF.org on 2024-11-02. 2023.</mixed-citation></citation-alternatives></ref><ref id="cit29"><label>29</label><citation-alternatives><mixed-citation xml:lang="ru">Mace G.M., Collar N.J., Gaston K.J., et al. Quantification of extinction risk: IUCN’s System for Classifying Threatened Species // Conservation Biology. 2008. Vol. 22. (6). P. 1424-1442. DOI: 10.1111/j.1523-1739.2008.01044.x.</mixed-citation><mixed-citation xml:lang="en">Mace G.M., Collar N.J., Gaston K.J., et al. Quantification of extinction risk: IUCN’s System for Classifying Threatened Species // Conservation Biology. 2008. Vol. 22. (6). P. 1424-1442. DOI: 10.1111/j.1523-1739.2008.01044.x.</mixed-citation></citation-alternatives></ref><ref id="cit30"><label>30</label><citation-alternatives><mixed-citation xml:lang="ru">Miller R.M., Rodriguez J.P., Aniskowicz-Fowler T., et al. National Threatened Species Listing based on IUCN Criteria and Regional Guidelines: Current status and future perspectives // Conservation Biology. 2007. Vol. 21 (3). P. 684-696. DOI: 10.1111/j.1523-1739.2007.00656.x.</mixed-citation><mixed-citation xml:lang="en">Miller R.M., Rodriguez J.P., Aniskowicz-Fowler T., et al. National Threatened Species Listing based on IUCN Criteria and Regional Guidelines: Current status and future perspectives // Conservation Biology. 2007. Vol. 21 (3). P. 684-696. DOI: 10.1111/j.1523-1739.2007.00656.x.</mixed-citation></citation-alternatives></ref><ref id="cit31"><label>31</label><citation-alternatives><mixed-citation xml:lang="ru">Norman Myers, Russell A. Mittermeier, Cristina G. Mittermeier, Gustavo A. B. da Fonseca &amp; Jennifer Kent. Biodiversity hotspots for conservation priorities // Nature. 2000. Vol. 403. P. 853-858.</mixed-citation><mixed-citation xml:lang="en">Norman Myers, Russell A. Mittermeier, Cristina G. Mittermeier, Gustavo A. B. da Fonseca &amp; Jennifer Kent. Biodiversity hotspots for conservation priorities // Nature. 2000. Vol. 403. P. 853-858.</mixed-citation></citation-alternatives></ref><ref id="cit32"><label>32</label><citation-alternatives><mixed-citation xml:lang="ru">Pham Hoang Ho. An illustrated flora of Vietnam // Monreal. 1991. Vol. 1. № 1. P. 42-261.</mixed-citation><mixed-citation xml:lang="en">Pham Hoang Ho. An illustrated flora of Vietnam // Monreal. 1991. Vol. 1. № 1. P. 42-261.</mixed-citation></citation-alternatives></ref><ref id="cit33"><label>33</label><citation-alternatives><mixed-citation xml:lang="ru">Pham V.V., Ammer C., Annighofer P. The presence of IUCN Red List Tree species in dependence of site characteristics in the Vietnamese Cat Ba National Park // Diversity. 2020. Vol. 12. № 3. Р. 104. DOI: 10.3390/d12030104.</mixed-citation><mixed-citation xml:lang="en">Pham V.V., Ammer C., Annighofer P. The presence of IUCN Red List Tree species in dependence of site characteristics in the Vietnamese Cat Ba National Park // Diversity. 2020. Vol. 12. № 3. Р. 104. DOI: 10.3390/d12030104.</mixed-citation></citation-alternatives></ref><ref id="cit34"><label>34</label><citation-alternatives><mixed-citation xml:lang="ru">Phan Ke Loc. The updated checklist of the fern flora of Vietnam following the classification scheme of A. Smith et al. (2006) // Journal of Fairylake Botanical Garden. 2010. Vol. 9. №3. P. 1-14.</mixed-citation><mixed-citation xml:lang="en">Phan Ke Loc. The updated checklist of the fern flora of Vietnam following the classification scheme of A. Smith et al. (2006) // Journal of Fairylake Botanical Garden. 2010. Vol. 9. №3. P. 1-14.</mixed-citation></citation-alternatives></ref><ref id="cit35"><label>35</label><citation-alternatives><mixed-citation xml:lang="ru">Polypodiopsida in GBIF Secretariat. GBIF Backbone Taxonomy. Checklist dataset doi: 10.15468/39omei accessed via GBIF.org on 2024-11-02. 2023.</mixed-citation><mixed-citation xml:lang="en">Polypodiopsida in GBIF Secretariat. GBIF Backbone Taxonomy. Checklist dataset doi: 10.15468/39omei accessed via GBIF.org on 2024-11-02. 2023.</mixed-citation></citation-alternatives></ref><ref id="cit36"><label>36</label><citation-alternatives><mixed-citation xml:lang="ru">Possingham H., Andelman S., Burgman M.A., et al. Limits to the use of threatened species lists // Trends in Ecology &amp; Evolution. 2002. Vol. 17. № 11. P. 503–507. DOI: 10.1016/S0169-5347(02)02614-9</mixed-citation><mixed-citation xml:lang="en">Possingham H., Andelman S., Burgman M.A., et al. Limits to the use of threatened species lists // Trends in Ecology &amp; Evolution. 2002. Vol. 17. № 11. P. 503–507. DOI: 10.1016/S0169-5347(02)02614-9</mixed-citation></citation-alternatives></ref><ref id="cit37"><label>37</label><citation-alternatives><mixed-citation xml:lang="ru">Rodrigues A., Pilgrim J., Lamoreux J., et al. The value of the IUCN Red List for conservation // Trends in Ecology &amp; Evolution. 2006. Vol. 21. № 2. 71-76. DOI: 10.1016/j.tree.2005.10.010</mixed-citation><mixed-citation xml:lang="en">Rodrigues A., Pilgrim J., Lamoreux J., et al. The value of the IUCN Red List for conservation // Trends in Ecology &amp; Evolution. 2006. Vol. 21. № 2. 71-76. DOI: 10.1016/j.tree.2005.10.010</mixed-citation></citation-alternatives></ref><ref id="cit38"><label>38</label><citation-alternatives><mixed-citation xml:lang="ru">The Biodiversity Law: 20/2008/QH12, 2008, [online] Available at http://portal.gms-eoc.org/uploads/resources/953/attachment/Vietname_Law_on_Boidiversity_28_Nov_2008_En.pdf.</mixed-citation><mixed-citation xml:lang="en">The Biodiversity Law: 20/2008/QH12, 2008, [online] Available at http://portal.gms-eoc.org/uploads/resources/953/attachment/Vietname_Law_on_Boidiversity_28_Nov_2008_En.pdf.</mixed-citation></citation-alternatives></ref><ref id="cit39"><label>39</label><citation-alternatives><mixed-citation xml:lang="ru">The Rio Declaration on Environment and Development United Nations. General Assembly Report of the United Nations Conference on Environment and Development* - Annex 1. Rio de Janeiro. 1992.</mixed-citation><mixed-citation xml:lang="en">The Rio Declaration on Environment and Development United Nations. General Assembly Report of the United Nations Conference on Environment and Development* - Annex 1. Rio de Janeiro. 1992.</mixed-citation></citation-alternatives></ref><ref id="cit40"><label>40</label><citation-alternatives><mixed-citation xml:lang="ru">Thin. N.N. Biodiversity and resources of plant heredity. National University Press, 2000. Ha Noi.</mixed-citation><mixed-citation xml:lang="en">Thin. N.N. Biodiversity and resources of plant heredity. National University Press, 2000. Ha Noi.</mixed-citation></citation-alternatives></ref><ref id="cit41"><label>41</label><citation-alternatives><mixed-citation xml:lang="ru">Hai He, Liang Zhang, Xin-Mao Zhou, Xin-Fen Gao and Li-Bing Zhang. Using a multilocus phylogeny to test morphology-based classifications of Polystichum (Dryopteridaceae), one of the largest fern genera // BMC Evolutionary Biology. 2016. Vol. 16, 55. DOI: 10.1186/s12862-016-0626-z.</mixed-citation><mixed-citation xml:lang="en">Hai He, Liang Zhang, Xin-Mao Zhou, Xin-Fen Gao and Li-Bing Zhang. Using a multilocus phylogeny to test morphology-based classifications of Polystichum (Dryopteridaceae), one of the largest fern genera // BMC Evolutionary Biology. 2016. Vol. 16, 55. DOI: 10.1186/s12862-016-0626-z.</mixed-citation></citation-alternatives></ref><ref id="cit42"><label>42</label><citation-alternatives><mixed-citation xml:lang="ru">Tran P., Tran T., Le D., et al. Vietnam Red Data Book - Part II. Plants. Vietnam Red Data Book. Vietnam Academy of Science and Technology Publishing House: Hanoi, 2007. 691 p.</mixed-citation><mixed-citation xml:lang="en">Tran P., Tran T., Le D., et al. Vietnam Red Data Book - Part II. Plants. Vietnam Red Data Book. Vietnam Academy of Science and Technology Publishing House: Hanoi, 2007. 691 p.</mixed-citation></citation-alternatives></ref><ref id="cit43"><label>43</label><citation-alternatives><mixed-citation xml:lang="ru">UNDP, SCBD, UNEP-WCMC. Creating a nature-positive future: The contribution of protected areas and other effective area-based conservation measures. UNDP. New York, NY. https://www.undp.org/sites/g/files/zskgke326/files/2021-11/UNDP-UNEP-Creating-a-Nature-Positive-Future-EN.pdf. 2021.</mixed-citation><mixed-citation xml:lang="en">UNDP, SCBD, UNEP-WCMC. Creating a nature-positive future: The contribution of protected areas and other effective area-based conservation measures. UNDP. New York, NY. https://www.undp.org/sites/g/files/zskgke326/files/2021-11/UNDP-UNEP-Creating-a-Nature-Positive-Future-EN.pdf. 2021.</mixed-citation></citation-alternatives></ref><ref id="cit44"><label>44</label><citation-alternatives><mixed-citation xml:lang="ru">UNEP-WCMC. Protected Planet Live Report 2023. www.protectedplanet.net [accessed Sept. 2023].</mixed-citation><mixed-citation xml:lang="en">UNEP-WCMC. Protected Planet Live Report 2023. www.protectedplanet.net [accessed Sept. 2023].</mixed-citation></citation-alternatives></ref><ref id="cit45"><label>45</label><citation-alternatives><mixed-citation xml:lang="ru">Vu Nguen Tu. New species of the genus Microsorum Link. From Vietnam // Botanical Journal. 1979. Vol. 64. № 12. P. 1769-1773.</mixed-citation><mixed-citation xml:lang="en">Vu Nguen Tu. New species of the genus Microsorum Link. From Vietnam // Botanical Journal. 1979. Vol. 64. № 12. P. 1769-1773.</mixed-citation></citation-alternatives></ref><ref id="cit46"><label>46</label><citation-alternatives><mixed-citation xml:lang="ru">Almond R.E.A., Grooten M., Petersen T. WWF. Living Planet Report 2020 Bending the curve of biodiversity loss. Switzerland: Gland, 2020. 164 p</mixed-citation><mixed-citation xml:lang="en">Almond R.E.A., Grooten M., Petersen T. WWF. Living Planet Report 2020 Bending the curve of biodiversity loss. Switzerland: Gland, 2020. 164 p</mixed-citation></citation-alternatives></ref><ref id="cit47"><label>47</label><citation-alternatives><mixed-citation xml:lang="ru">Xu H, Cao Y, Yu D, et al. Ensuring effective implementation of the post-2020 global biodiversity targets // Nature Ecology &amp; Evolution. 2021. Vol. 5. № 4. P. 411-418. DOI: 10.1038/s41559-020-01375-y</mixed-citation><mixed-citation xml:lang="en">Xu H, Cao Y, Yu D, et al. Ensuring effective implementation of the post-2020 global biodiversity targets // Nature Ecology &amp; Evolution. 2021. Vol. 5. № 4. P. 411-418. DOI: 10.1038/s41559-020-01375-y</mixed-citation></citation-alternatives></ref></ref-list><fn-group><fn fn-type="conflict"><p>The authors declare that there are no conflicts of interest present.</p></fn></fn-group></back></article>
